John Freeman

Associate Department Chair
Dewey B. and Velma P. Stuit Professor

John Freeman CV

Research Interests

Behavioral neuroscience; neuroscience of learning; categorization; comparative cognition 

Training Areas

Behavioral & Cognitive Neuroscience

Representative Publications

Broschard, M.B., Kim, J., Love, B.C., Wasserman, E.A., & Freeman, J.H. (2021). Prelimbic cortex maintains attention to category-relevant information and flexibly updates category representations. Neurobiology of Learning and Memory, 185, doi: 10.1016/j.nlm.2021.107524.

O’Donoghue, E.M., Broschard, M.B., Freeman, J.H., & Wasserman, E.A. (2022). The lords of the rings: People and pigeons take different paths mastering the concentric-rings categorization task. Cognition, 218, doi: 10.1016/j.cognition.2021.104920.

Broschard, M.B., Kim, J., Love, B.C., & Freeman, J.H. (2023). Dorsomedial striatum, but not dorsolateral striatum, is necessary for rat category learning. Neurobiology of Learning and Memory, 199, doi: 10.1016/j.nlm.2023.107732.

Halverson, H.E., Kim, J., & Freeman, J.H. (2023). Dynamic changes in local activity and network interactions among the anterior cingulate, amygdala and cerebellum during associative learning. Journal of Neuroscience, 43, 8385-8402.

Research areas
  • Behavioral Neuroscience
  • Categorization
  • Development
  • Learning and Memory
  • Neural Plasticity
john freeman.jpeg
Ph.D., University of North Carolina, 1994

379 Psychological and Brain Sciences Building (PBSB)
Iowa City, IA 52242
United States


307 Spence Labs (SLP)
Iowa City, IA 52242
United States

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